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Abstract Water quality and freshwater ecosystems are affected by river discharge and temperature. Models are frequently used to estimate river temperature on large spatial and temporal scales due to limited observations of discharge and temperature. In this study, we use physically based river routing and temperature models to simulate daily discharge and river temperature for rivers in 138 basins in Alaska, including the entire Yukon River basin, from 1990–2021. The river temperature model was optimized for ice free months using a surrogate‐based model optimization method, improving model performance at uncalibrated river gages. A common statistical model relating local air and water temperature was used as a benchmark. The physically based river temperature model exhibited superior performance compared to the benchmark statistical model after optimization, suggesting river temperature model optimization could become more routine. The river temperature model demonstrated high sensitivity to air temperature and model parameterization, and lower sensitivity to discharge. Validation of the models showed a Kling‐Gupta Efficiency of 0.46 for daily river discharge and a root mean square error of 2.04°C for daily river temperature, improving on the non‐optimized physical model and the benchmark statistical model, which had root mean square errors of 3.24 and 2.97°C, respectively. The simulation shows that rivers in northern Alaska have higher maximum summer temperatures and more variability than rivers in the Central and Southern regions. Furthermore, this framework can be readily adapted for use across models and regions. 

Abstract Record high temperatures were documented in the McMurdo Dry Valleys, Antarctica, on 18 March 2022, exceeding average temperatures for that day by nearly 30°C. Satellite imagery and stream gage measurements indicate that surface wetting coincided with this warming more than 2 months after peak summer thaw and likely exceeded thresholds for rehydration and activation of resident organisms that typically survive the cold and dry conditions of the polar fall in a freeze‐dried state. This weather event is notable in both the timing and magnitude of the warming and wetting when temperatures exceeded 0°C at a time when biological communities and streams have typically entered a persistent frozen state. Such events may be a harbinger of future climate conditions characterized by warmer temperatures and greater thaw in this region of Antarctica, which could influence the distribution, activity, and abundance of sentinel taxa. Here we describe the ecosystem responses to this weather anomaly reporting on meteorological and hydrological measurements across the region and on later biological observations from Canada Stream, one of the most diverse and productive ecosystems within the McMurdo Dry Valleys.